shaft,engine mounts,wishbones can be ..
but imo you must feel some shudder at the steering also (not like balance problem)
edit: wishbones and mounts can be checked simply by a lever but having witnessed shaft balance , they check it off the ground and you needto close the tc..
did you try full geometry check before ?
Hello Cem, nice of you to answer so quickly 
I can hardly feel the vibration through the steering wheel at all, it is through the accellerator pedal & car body. I haven't had the time yet to find a place relatively close to me that does full geometry work, as WIM is too far South for me to travel at the moment. I will ask my son to check the suspension linkages ASAP.
vibrations not easy to find if its not a geometry problem.. :-/
being sensitive , I spend considerable time and money for that..
for you to pinpoint the problem try using high tire pressures (for them not to compansate), this time you will feel vibrations more clear..and see at what speed they begin..
you said
" I have also noticed the shudder decreases slightly when I go downhill & increases uphill, so must be associated with engine load"
although this lessens the probability of geometry problem, personally I would still start from there .. to eliminate and be sure about the causes one by one..
edit:
also my experience says engine mounts and wishbones must start vibration earlier .. so let someone drive and you sit behind for control..